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Biogeography
of the Earth
The Alpine Tundra
The tundra biome is found at high elevations in mountainous terrain as
well. Many, if not all, the same features of the arctic tundra are found in the
alpine tundra. Microclimate is an important control over the distribution of plant species as
slope and exposure control the availability of light and moisture. The landscape is dotted
with small cushion plants, lichens and mosses. Willows are found where moisture is
abundant. Other factors like soil development, movement of soil by animals, and drainage
determine vegetation community distribution.
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Figure
BE.50 tundra vegetation (Photo credit: Michael Ritter)
Vegetation
consists of low growing shrubs, cushion plants, small forbs exploding with colorful
flowers and lush meadows of sedges and grasses. These plants cover gentle slopes and rock
crevices. Rock surfaces are dotted with a cover of lichens and mosses. Most species are
slow-growing perennials. Plants have been forced to adapt to such an extreme environment.
Ninety percent of total structure in some plants is in roots storing nutrients and energy
during poor growing periods. Flowers are often large but other parts of the plant are
small to save energy, and reducing exposure to the rigors of the wind. Some plants have
waxy coatings or hairs thus losing minimal heat and water to the wind. The location of
plant communities is correlated with the duration of snow cover. While snow is blown free
from exposed sites, it accumulates in the lee of obstructions and in depressions.
Community location is also related to soil, drainage, and movement of soil by burrowing
animals, and frost action which is prevalent throughout much of the alpine tundra. Dense
willow thickets often occupy moist depressions on the lee side of ridges. A deep cover of
snow during the winter protects buds from the wind and freezing temperatures. These are
the tallest perennials growing above the krummholz of the ecotone. 
Figure BE.51 Alpine
tundra fell field (foreground) and on rocky knoll ( upper right)
Colorado, USA. (Courtesy UNESCO)
Fell fields are colorful rock gardens exposed to the rigors of the wind. Wind
removes snow throughout the season subjecting the plants to desiccation. Low-lying mats
and cushion plants nestle against the rocky surface.
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Figure
BE.52 The alpine tundra of Niwot Ridge in early May
(Photo credit: Michael
Ritter)
In the Indian Peaks region of Colorado, the alpine tundra ranges from
11,200 to 12,000 feet depending on latitude and slope exposure. Plant communities vary
significantly in shape and plant composition, and may vary in size from a few square
inches to several acres.
The climate of the tundra is exceedingly harsh. Annual precipitation is around 40
inches, effective precipitation is far below that amount however. Snow remains as
permanent snow fields at some sites. Wind speeds can exceed 100 mph and mean annual
temperature is below freezing. The frost free season approx. 1 1/2 months. Diurnal
temperature ranges are small because the air is mixed by the constant winds.
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Figure BE.53
Solifluction terraces
with snow lying
behind. (Photo credit: Michael Ritter)
Soils are quite variable, from barely any soils in valleys scoured by glaciers to the
mature residual soils of unglaciated ridges, and scattered in between rocks brought to
surface from frost heave to form periglacial features like polygons. Soil ice is
found in all soils in winter, and soil temperatures are low enough to form patches of
permafrost. A common landscape feature of the tundra are solifluction
terraces. Solifluction terraces with willow growing on top and to the back are
visible in the center of the photograph. Late lying snow patches keep conditions moist to
permit willow growth. Reaching a few feet high, willows are covered by snow to protect
the over-wintering buds. Willows are the tallest of any species in the alpine tundra.
Figure
BE.54 Polygon, outlined in black, caused by frost heave. (Photo credit:
Michael Ritter)
The plant communities mentioned above are considered climax communities mainly because
they change so slowly. Communities are often disturbed by small burrowing animals like the
pocket gopher that churn up the soil and eat plant roots, or voles which can devastate
above - ground biomass. Recovery after disturbance proceeds exceedingly slow, slower than
any other mountain ecosystem.
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